Dinosaurs Live Misc Note 2: Dinosaurs featured in Power Rangers Dino Thunder

The Zords of Power Rangers Dino Thunder were dinosaur-based cyborgs created by Tommy Oliver and Anton Mercer during their collaboration. They were mostly maintained by Tommy's friend Hayley. The Dinozords (first called Biozords), created by Anton Mercer and Tommy Oliver with the rest of their dinosaur-related experiments, they were commandeered by Mesogog. He unleashed the Tyrannozord, Pterazord, and Tricerazord upon the Rangers, but they were taken over by the Dino Thunder Rangers. Other Dinozords were unleashed throughout the series. The Tyrannozord, Tricerazord, Pterazord, and the Mezodon Rover were destroyed battling the Zelzord. The fate of the other Zords was not shown or discussed on screen.

Primary Zords
Tyrannozord: A red Tyrannosaurus Rex Zord that forms the majority of the Thundersaurus Megazord (head, torso, left arm, legs and the Dino Drill), controlled by Conner McKnight the Red Dino Ranger. It was destroyed in the finale.
Tricerazord: A blue Triceratops Zord that forms the right arm and Toro Shield of the Thundersaurus Megazord, controlled by Ethan James the Blue Dino Ranger. It was destroyed in the finale.
Pterazord: A yellow Pteranodon Zord that forms helmet and chestplate of the Thundersaurus Megazord, as well at the Pterarang weapon, controlled by Kira Ford the Yellow Dino Ranger. It was destroyed in the finale.
Brachiozord: A black Brachiosaurus Carrierzord that carries the Tyranno, Tricera, and Ptera Zords, as well as the Auxiliary Zords. Has been shown to fire electric blasts from its mouth. Controlled by Tommy Oliver the Black Dino Ranger. This may be a reference to Titanus the Carrierzord from Mighty Morphin Power Rangers.
Dragozord: A white Tupuxuara Zord that forms arms, legs, and weapons of the Dino Stegazord, and the wings of the Valkasaurus Megazord. Can create gusts of wind by flapping its wings. Controlled by Trent Fernandez the White Dino Ranger. Where it got its name is a mystery since "Drago" means Dragon in Latin.
Mezodon Rover: The personal Zord of the Triassic Ranger, the Mezodon Rover has two forms. One is that of a giant chariot pulled by the "Styracozord" (a crimson colored Styracosaurus zord), called the Triassic Megarover. In the chariot form, the Mezodon Rover's wheels can ignite and be used to ram into enemies. It also was controlled by Tommy (in 'A Test of Trust') and Ethan (in 'Thunder Struck'). It can combine with Cephalozord, Dimetrazord, Parasaurzord and Ankylozord to form Triceramax Megazord. It was destroyed by Ethan to destroy the Zelzord after it was set to self destruct.

Auxiliary Zords
Cephalozord: A violet Pachycephalosaurus Zord that forms the Punch weapon arm (right side) for the Thundersaurus Megazord and Dino Stegazord. It also forms the right arm of the Triceramax Megazord. Usually summoned by Tommy, but can also be controlled by Ethan.
Dimetrozord: A cyan Dimetrodon Zord that forms the Saw weapon arm (left side) for the Thundersaurus Megazord and Dino Stegazord. It also forms the inner right leg of the Triceramax Megazord. Usually summoned by Tommy, but can also be controlled by Kira.
Stegozord: A crimson Stegosaurus Zord that becomes a hovercraft for the Thundersaurus and Valkasaurus Megazord. Helps the Thundersaurus Megazord do a "Surf's Up" slash/drill attack. It forms the body and the head for the Dino Stegazord. Traditionally, it is Trent's Zord but it could also be Tommy's second Zord, seeing as how he entered Dino Stegazord, when he temporarily commanded it, and the Valkasaurus Megazord through the Stegozord.
Parasaurzord: A green Parasaurolophus Zord that forms the Scissor Blade weapon (left side) arm for the Thundersaurus Megazord. It also forms the inner left leg of the Triceramax Megazord. Usually summoned by Tommy, but can be controlled by Kira.
Ankylozord: An orange Ankylosaurus Zord that forms the Shield Drill weapon arm (right side) for the Thundersaurus Megazord. It also forms the left arm of the Triceramax Megazord. Usually summoned by Tommy, but can also be controlled by Ethan.

Replicant Zords
Carnotaur Zord: An indigo-and-teal Carnotaurus Zord that forms the majority of the Blizzard Megazord (head, chest, legs, left arm, and the Blizzard Drill).
Chasmosaur Zord: A silver-and-violet Chasmosaurus Zord that forms the right arm and the Blizzard Shield of the Blizzard Megazord.

Dinosaurs Live highlight 28: Triceratops

Triceratops is a genus of herbivorous ceratopsid dinosaur which lived during the late Maastrichtian stage of the Late Cretaceous Period, around 68 to 65 million years ago (Mya) in what is now North America. It was one of the last dinosaur genera to appear before the great Cretaceous–Paleogene extinction event. The term Triceratops, which literally means "three-horned face," is derived from the Greek τρί- (tri-) meaning "three", κέρας (kéras) meaning "horn", and ὤψ (ops) meaning "face".

Bearing a large bony frill and three horns on its large four-legged body, and conjuring similarities with the modern rhinoceros, Triceratops is one of the most recognizable of all dinosaurs and the best known ceratopsid. It shared the landscape with and was preyed upon by the fearsome Tyrannosaurus, though it is less certain that the two did battle in the manner often depicted in traditional museum displays and popular images.

The exact placement of the Triceratops genus within the ceratopsid group has been debated by paleontologists. Two species, T. horridus and T. prorsus, are considered valid although many other species have been named. Recent research suggests that the contemporaneous Torosaurus, a ceratopsid long regarded as a separate genus, actually represents Triceratops in its mature form.

Triceratops has been documented by numerous remains collected since the genus was first described in 1889, including at least one complete individual skeleton. Paleontologist John Scannella observed: "It is hard to walk out into the Hell Creek Formation and not stumble upon a Triceratops weathering out of a hillside." Forty-seven complete or partial skulls were discovered in just that area during the decade 2000–2010. Specimens representing life stages from hatchling to adult have been found.

Description
The function of the frills and three distinctive facial horns has long inspired debate. Traditionally these have been viewed as defensive weapons against predators. More recent theories, noting the presence of blood vessels in the skull bones of ceratopsids, find it more probable that these features were primarily used in identification, courtship and dominance displays, much like the antlers and horns of modern reindeer, mountain goats, or rhinoceros beetles. The theory finds additional support if Torosaurus represents the mature form of Triceratops, as this would mean the frill also developed holes (fenestrae) as individuals reached maturity, rendering the structure more useful for display than defense.

Individual Triceratops are estimated to have reached about 7.9 to 9.0 m (26.0–29.5 ft) in length, 2.9 to 3.0 m (9.5–9.8 ft) in height, and 6.1–12.0 tonnes (13,000–26,000 lb) in weight. The most distinctive feature is their large skull, among the largest of all land animals. The largest known skull (specimen BYU 12183) is estimated to have been 2.5 metres (8.2 ft) in length when complete, and could reach almost a third of the length of the entire animal. It bore a single horn on the snout, above the nostrils, and a pair of horns approximately 1 m (3 ft) long, with one above each eye. To the rear of the skull was a relatively short, bony frill, adorned with epoccipitals in some specimens. Most other ceratopsids had large fenestrae in their frills, while those of Triceratops were noticeably solid.

Dinosaurs Live highlight 27: Stegosaurus

Stegosaurus (meaning "roof lizard" or "covered lizard" in reference to its bony plates) is a genus of armored stegosaurid dinosaur. They lived during the Late Jurassic period (Kimmeridgian to early Tithonian), some 155 to 150 million years ago in what is now western North America. In 2006, a specimen of Stegosaurus was announced from Portugal, showing that they were present in Europe as well. Due to its distinctive tail spikes and plates, Stegosaurus is one of the most recognizable dinosaurs. At least three species have been identified in the upper Morrison Formation and are known from the remains of about 80 individuals.

A large, heavily built, herbivorous quadruped, Stegosaurus had a distinctive and unusual posture, with a heavily rounded back, short forelimbs, head held low to the ground and a stiffened tail held high in the air. Its array of plates and spikes has been the subject of much speculation. The spikes were most likely used for defense, while the plates have also been proposed as a defensive mechanism, as well as having display and thermoregulatory functions. Stegosaurus had a relatively low brain-to-body mass ratio. It had a short neck and small head, meaning it most likely ate low-lying bushes and shrubs. It was the largest of all the stegosaurians (bigger than genera such as Kentrosaurus and Huayangosaurus) and, although roughly bus-sized, it nonetheless shared many anatomical features (including the tail spines and plates) with the other stegosaurian genera.

Description
Averaging around 9 meters (30 ft) long and 4 meters (14 ft) tall, the quadrupedal Stegosaurus is one of the most easily identifiable dinosaurs, due to the distinctive double row of kite-shaped plates rising vertically along its rounded back and the two pairs of long spikes extending horizontally near the end of its tail. Although a large animal, it was dwarfed by its contemporaries, the giant sauropods. Some form of armor appears to have been necessary, as it coexisted with large predatory theropod dinosaurs, such as the fearsome Allosaurus and Ceratosaurus.

The hind feet each had three short toes, while each forefoot had five toes; only the inner two toes had a blunt hoof. All four limbs were supported by pads behind the toes. The forelimbs were much shorter than the stocky hindlimbs, which resulted in an unusual posture. The tail appears to have been held well clear of the ground, while the head of Stegosaurus was positioned relatively low down, probably no higher than 1 meter (3.3 ft) above the ground.

The long and narrow skull was small in proportion to the body. It had a small antorbital fenestra, the hole between the nose and eye common to most archosaurs, including modern birds, though lost in extant crocodylians. The skull's low position suggests that Stegosaurus may have been a browser of low-growing vegetation. This interpretation is supported by the absence of front teeth and their replacement by a horny beak or rhamphotheca. Stegosaurian teeth were small, triangular and flat; wear facets show that they did grind their food. The inset placement in the jaws suggests that Stegosaurus had cheeks to keep food in their mouths while they chewed.

Despite the animal's overall size, the braincase of Stegosaurus was small, being no larger than that of a dog. A well-preserved Stegosaurus braincase allowed Othniel Charles Marsh to obtain in the 1880s a cast of the brain cavity or endocast of the animal, which gave an indication of the brain size. The endocast showed that the brain was indeed very small, maybe the smallest among the dinosaurs. The fact that an animal weighing over 4.5 metric tons (5 short tons) could have a brain of no more than 80 grams (2.8 oz) contributed to the popular old idea that dinosaurs were unintelligent, an idea now largely rejected.

Dinosaurs Live highlight 26: Ornithomimus

Ornithomimus ("bird mimic") is a genus of ornithomimid dinosaur from the Late Cretaceous Period of what is now North America.

In 1890 Ornithomimus velox was named by Othniel Charles Marsh on the basis of a foot and partial hand from the Maastrichtian Denver Formation. Another seventeen species have been named since. Most of these have subsequently been assigned to new genera or shown to be not directly related to Ornithomimus. The best material of species still considered part of the genus has been found in Canada, representing the earlier Edmontonian-age Ornithomimus edmontonicus Sternberg 1933, known from several skeletons. However, on some of these the new genus Dromiceiomimus including Dromiceiomimus brevitertius (Parks 1926) has been based, causing taxonomic problems of priority and identity that are still unresolved.

Ornithomimus was a relatively small swift bipedal animal, equipped with a small toothless beaked head, that may indicate an omnivorous diet.

Description
Like other ornithomimids, Ornithomimus is characterized by a foot with three weight-bearing toes, long slender arms and a long neck with a birdlike, elongated, toothless, beaked skull. It was bipedal and superficially resembled an ostrich, except for its long tail. It would have been a swift runner. It had very long limbs, hollow bones, and a large brain and eyes. The brains of ornithomimids were large for dinosaurs, but this may not necessarily be a sign of greater intelligence; some paleontologists think that the enlarged portions of the brain were dedicated to kinesthetic coordination. Their hands are remarkably sloth-like in appearance, which led Henry Fairfield Osborn to suggest that they were used to hook branches during feeding.

Ornithomimus differs from other ornithomimids, such as Struthiomimus, in having a short back, long slender forearms, very slender, straight hand and foot claws and in having metacarpals and fingers of similar lengths.

The three Ornithomimus species today seen as possibly valid, differ rather in size. In 2010 Gregory S. Paul estimated the length of O. edmontonicus at 3.8 metres, its weight at 170 kilograms (370 lb). One of its specimens, CMN 12228, preserves a femur (thigh bone) 46.8 centimetres (18.4 in) long. O. sedens was by Paul estimated at 4.8 metres and 350 kilograms (770 lb). O. velox, the type species of Ornithomimus, is based on material of a much smaller animal. Whereas the holotype of O. edmontonicus, CMN 8632, preserves a second metacarpal eighty-four millimetres long, the same element with O. velox measures only fifty-three millimetres.

Paleobiology
The diet of Ornithomimus is still debated. As theropods, ornithomimids might have been carnivorous but their body shape would also have been suited for a partly or largely herbivorous lifestyle. Suggested food includes insects, crustaceans, fruit, leaves, branches, eggs, and the meat of lizards and small mammals.

Ornithomimus had legs that seem clearly suited for rapid locomotion, with the tibia about 20% longer than the femur. The large eye sockets suggest a keen visual sense, and also suggest the possibility that they were nocturnal.

Dinosaurs Live highlight 25: Iguanodon

Iguanodon (meaning "iguana-tooth") is a genus of ornithopod dinosaur that lived roughly halfway between the first of the swift bipedal hypsilophodontids and the ornithopods' culmination in the duck-billed dinosaurs. Many species of Iguanodon have been named, dating from the Kimmeridgian age of the Late Jurassic Period to the Cenomanian age of the Late Cretaceous Period from Asia, Europe, and North America. However, research in the first decade of the 21st century suggests that there is only one well-substantiated species: I. bernissartensis, that lived from the late Barremian to the earliest Aptian ages (Early Cretaceous) in Belgium, between about 126 and 125 million years ago. Iguanodon's most distinctive features were its large thumb spikes, which were possibly used for defence against predators, combined with long prehensile fifth fingers able to forage for food.

Named in 1825 by English geologist Gideon Mantell, Iguanodon was the second dinosaur formally named, after Megalosaurus. Together with Megalosaurus and Hylaeosaurus, it was one of the three genera originally used to define Dinosauria. A large, bulky herbivore, Iguanodon is a member of Iguanodontia, along with the duck-billed hadrosaurs. The taxonomy of this genus continues to be a topic of study as new species are named or long-standing ones reassigned to other genera.

Scientific understanding of Iguanodon has evolved over time as new information has been obtained from the fossils. The numerous specimens of this genus, including nearly complete skeletons from two well-known bonebeds, have allowed researchers to make informed hypotheses regarding many aspects of the living animal, including feeding, movement, and social behaviour. As one of the first scientifically well-known dinosaurs, Iguanodon has occupied a small but notable place in the public's perception of dinosaurs, its artistic representation changing significantly in response to new interpretations of its remains.

Description
Iguanodon was a bulky herbivore that could shift from bipedality to quadrupedality. The best-known species, I. bernissartensis, is estimated to have weighed about 3 tonnes (3.5 tons) on average, and measured about 10 metres long (33 ft) as an adult, with some specimens possibly as long as 13 metres (43 ft). This genus had a large, tall but narrow skull, with a toothless beak probably covered with keratin, and teeth like those of an iguana, but much larger and more closely packed.

The arms were long (up to 75% the length of the legs in I. bernissartensis) and robust,[4] with rather inflexible hands built so that the three central fingers could bear weight. The thumbs were conical spikes that stuck out away from the three main digits. In early restorations, the spike was placed on the animal's nose. Later fossils revealed the true nature of the thumb spikes, although their exact function is still debated. They could have been used for defense, or for foraging for food. The little finger was elongated and dextrous, and could have been used to manipulate objects. The legs were powerful, but not built for running, and each foot had three toes. The backbone and tail were supported and stiffened by ossified tendons, which were tendons that turned to bone during life (these rod-like bones are usually omitted from skeletal mounts and drawings). Overall, in body structure, it was not too dissimilar from its later relatives, the hadrosaurids.

Dinosaurs Live highlight 24: Nanotyrannus

Nanotyrannus ("dwarf tyrant") is a genus of tyrannosaurid dinosaur, known only from two juvenile specimens, which may in fact represent juvenile specimens of the contemporary species Tyrannosaurus rex.

Description: Differences from the T-Rex
The primary differences that some scientists have used to distinguish Nanotyrannus lancensis from Tyrannosaurus rex primarily concern the number of teeth. Nanotyrannus had more teeth in its upper and lower jaws than an adult Tyrannosaurus. N. lancensis had 14-15 teeth in each side of the upper jaw (maxilla) and 16 teeth in each side of the lower jaw (dentary). T. rex, on the other hand, had 11-12 tooth positions in the upper jaw and 11-14 in the lower. The exact implications of this difference in tooth count has been controversial. In his 1999 study of tyrannosaurid growth patterns, Carr showed that in Gorgosaurus libratus, the number of teeth decreased as the animal grew, and he used this data to support the hypothesis that N. lancensis is simply a juvenile T. rex. The team of scientists who studied growth in the related Tarbosaurus bataar found little to no decrease in tooth count as that species grew, even though they had juvenile specimens much younger than the Nanotyrannus specimens. These researchers also noted, however, that in both Tyrannosaurus and Gorgosaurus, there were significant differences in tooth count between individuals of the same age group, and that tooth count may vary on an individual basis not related to growth.

Another difference cited by those who support the validity of N. lancensis is the presence of a small foramen, or pit, in the quadratojugal, a bone in the back corner of the skull. Both the holotype and the "Jane" specimen have this feature, suggesting it is not a deformity, and it is not known in any adult tyrannosaurid specimens. It is possible that this is again an individual variant, or that it was a feature lost as the animals grew, though studies of other juvenile tyrannosaurids do not show an equivalent feature.

Dinosaurs Live highlight 23: Utahraptor

Utahraptor (meaning "Utah's predator" or "Utah thief") is a genus of theropod dinosaurs, including the largest known members of the family Dromaeosauridae. Fossil specimens date to the upper Barremian stage of the early Cretaceous period (in rock strata dated to 126 ± 2.5 million years ago). It contains a single species, Utahraptor ostrommaysorum.

Description
The holotype specimen of Utahraptor is fragmentary, consisting of skull fragments, a tibia, claws and some caudal (tail) vertebra. These few elements suggest an animal about twice the size of Deinonychus. Like other dromaeosaurids, Utahraptor had large curved claws on their second toes. One claw specimen is preserved at 22 centimetres (8.7 in) in length and is thought to reach 24 centimetres (9.4 in) restored.

The largest described U. ostrommaysorum specimens are estimated to have reached up to 7 m (23 ft) long and somewhat less than 500 kg (1,100 lb) in weight, comparable to a grizzly bear in size. Some undescribed specimens in the BYU collections may have reached up to 11 m (36 ft) long, though these await more detailed study.

It is thought that Utahraptor may be closely related to the smaller Dromaeosaurus and the giant Mongolian dromaeosaurid genus Achillobator.

Although feathers have never been found in association with Utahraptor specimens, there is strong phylogenetic evidence suggesting that all dromaeosaurids possessed them. This evidence comes from phylogenetic bracketing, which allows paleontologists to infer traits that exist in a clade based on the existence of that trait in a more basal form. The genus Microraptor is one of the oldest known dromaeosaurids, and is phylogenetically more primitive than Utahraptor. Since Microraptor possessed feathers, it is reasonable to assume that this trait was present in all of Dromaeosauridae. Feathers were very unlikely to have evolved more than once in dromaeosaurids, so assuming that Utahraptor lacked feathers would require positive evidence that they did not have them. So far, there is nothing to suggest that feathers were lost in larger, more derived species of dromaeosaurs.

Dinosaurs Live highlight 22: Pachycephalosaurus

Pachycephalosaurus (meaning "thick headed lizard," from Greek pachys-/παχυς- "thick", kephale/κεφαλη "head" and sauros/σαυρος "lizard") is a genus of pachycephalosaurid dinosaur. It lived during the Late Cretaceous Period (Maastrichtian stage) of what is now North America. Remains have been excavated in Montana, South Dakota, and Wyoming. It was an herbivorous or omnivorous creature which is only known from a single skull and a few extremely thick skull roofs. This dinosaur is monotypic, meaning the type species, P. wyomingensis, is the only known species. Pachycephalosaurus was one of the last non-avian dinosaurs before the K–T extinction event. Another dinosaur, Tylosteus of western North America, has been synonymized with Pachycephalosaurus.

Like other pachycephalosaurids, Pachycephalosaurus was a bipedal omnivore with an extremely thick skull roof. It possessed long hindlimbs and small forelimbs. Pachycephalosaurus is the largest known pachycephalosaur.

The thick skull domes of Pachycephalosaurus and related genera gave rise to the hypothesis that pachycephalosaurs used their skulls in intraspecific combat. This hypothesis has been disputed in recent years.

Description
The anatomy of Pachycephalosaurus is poorly known, as only skull remains have been described. Pachycephalosaurus is famous for having a large, bony dome atop its skull, up to 25 cm (10 in) thick, which safely cushioned its tiny brain. The dome's rear aspect was edged with bony knobs and short bony spikes projected upwards from the snout. The spikes were probably blunt, not sharp.

The skull was short, and possessed large, rounded eye sockets that faced forward, suggesting that the animal had good vision and was capable of binocular vision. Pachycephalosaurus had a small muzzle which ended in a pointed beak. The teeth were tiny, with leaf-shaped crowns. The head was supported by an "S"- or "U"-shaped neck.

Pachycephalosaurus was probably bipedal and was the largest of the pachycephalosaurid (bone-headed) dinosaurs. It has been estimated that Pachycephalosaurus was around 4.5 metres (15 ft) long and weighed 450 kilograms (990 lb). Based on other pachycephalosaurids, it probably had a fairly short, thick neck, short fore limbs, a bulky body, long hind legs and a heavy tail, which was likely held rigid by ossified tendons.

Classification
Pachycephalosaurus gives its name to the Pachycephalosauria, a clade of herbivorous ornithischian ("bird hipped") dinosaurs which lived during the Late Cretaceous Period in North America and Asia. Despite their bipedal stance, they were likely more closely related to the ceratopsians than the ornithopods.

Dinosaurs Live highlight 21: Protoceratops

Protoceratops (from Greek proto-/πρωτο- "first", cerat-/κερατ- "horn" and -ops/-ωψ "face", meaning "First Horned Face") is a genus of sheep-sized (1.5 to 2 m long) herbivorous ceratopsian dinosaur, from the Upper Cretaceous Period (Campanian stage) of what is now Mongolia. It was a member of the Protoceratopsidae, a group of early horned dinosaurs. Unlike later ceratopsians, however, it was a much smaller creature that lacked well-developed horns and retained some primitive traits not seen in later genera.

Protoceratops had a large neck frill, which may have served to protect the neck, to anchor jaw muscles, to impress other members of the species, or combinations of these functions. Described by Walter Granger and W.K. Gregory in 1923, Protoceratops was initially believed to be an ancestor of the North American ceratopsians. Researchers currently distinguish two species of Protoceratops (P. andrewsi and P. hellenikorhinus), based in part by their respective sizes.

Description
Protoceratops was a quadrupedal dinosaur that was partially characterized by its distinctive neck frill at the back of its skull. The frill itself contained two large parietal fenestra (holes in the frill), while its cheeks had large jugal bones. The exact size and shape of the neck frill varied by individual; some specimens had short, compact frills, while others had frills nearly half the length of the skull. The frill consists mostly of the parietal bone and partially of the squamosal. Some researchers, including Peter Dodson attribute the different sizes and shapes of these bones to sexual dimorphism, as well as the age of the specimen, at the time of death.

Protoceratops was approximately 1.8 meters (6 ft) in length and 0.6 meters (2 ft) high at the shoulder. A fully grown adult would have weighed less than 400 pounds (180 kg). The large numbers of specimens found in high concentration suggest that Protoceratops lived in herds.

Protoceratops was a relatively small dinosaur with a proportionately large skull. Despite being herbivorous, Protoceratops appears to have had muscular jaws capable of a powerful bite. These jaws were packed with dozens of teeth, well suited for chewing tough vegetation. The skull consisted of a massive frontal beak, and four pairs of fenestrae (skull openings). The foremost hole, the naris, was considerably smaller than the nostrils seen in later genera. Protoceratops had large orbits (the holes for its eyes), which measured around 50 millimeters in diameter. Behind the eye was a slightly smaller fenestra, known as the "infratemporal fenestra."

Daily activity patterns
The large eyes of Protoceratops has been suggested as evidence for a nocturnal lifestyle. However, subsequent comparisons between the scleral rings of Protoceratops and modern birds and reptiles have indicated a more cathemeral lifestyle, being active throughout the day during short intervals. This suggests that the fight between Protoceratops and the primarily nocturnal Velociraptor indicated by the fighting specimens may have occurred at twilight or under low-light conditions.

Dinosaurs Live highlight 20: Omeisaurus

Omeisaurus (meaning "Omei lizard") is a genus of sauropod dinosaur from the Middle Jurassic Period (Bathonian-Callovian stage) of what is now China. Its name comes from Mount Emei, where it was discovered in the lower Shaximiao Formation of Sichuan Province.

Like other sauropods, Omeisaurus was herbivorous and large. It measured 10 to 15.2 metres (30 to 50 ft) long, 4 metres (12 ft) high and weighed 4 tons.

Omeisaurus was first described in 1939. It was named after the sacred mountain Omeishan, which is were the first fossil example of Omeisaurus was found. Most skeletons of Omeisaurus were found in the 1970s and 1980s, during the great “Chinese Dinosaur rush”. There have been six species of Omeisaurus named so far: O. junghsiensis, O. changshouensis, O. fuxiensis, O. tianfuensis, O. luoquanensis, and O. maoianus. All of these but the last were named after the locations where they were found. O. fuxiensis was the smallest species, measuring around 35 feet (11 m) long. O. tianfuensis had the longest neck of the genus, around 30 feet (9.1 m) long. The only dinosaur with a longer neck was Mamenchisaurus. A club tail fossil discovered in the same bone bed as the Omeisaurus fossils was thought to belong to this genus, but is now believed to belong to a large specimen of Shunosaurus.

Mounted skeletons of Omeisaurus are on display at the Zigong Dinosaur Museum in Zigong, Sichuan Province and at Beipei Museum, near Chongqing, both in China.

Dinosaurs Live highlight 19: Pteranodon

Pteranodon (from Greek πτερ- "wing" and αν-οδων "toothless"), from the Late Cretaceous geological period of North America in present day Kansas, Alabama, Nebraska, Wyoming, and South Dakota, was one of the largest pterosaur genera and had a maximum wingspan of over 6 metres (20 ft). Pteranodon is known from more fossil specimens than any other pterosaur, with about 1,200 specimens known to science, many of them well preserved, with complete skulls and articulated skeletons. It was an important genus of the animal community present in the Western Interior Seaway.

Pteranodon was a reptile, but not a dinosaur. By definition, all dinosaurs belong to the groups Saurischia and Ornithischia, which excludes pterosaurs. Nevertheless, Pteranodon is frequently featured in dinosaur books and is strongly associated with dinosaurs by the general public.

Desription
Pteranodon species are extremely well represented in the fossil record, allowing for detailed descriptions of their anatomy and analysis of their life history. Over 1,000 specimens have been identified, though less than half are complete enough to give researchers good information on the anatomy of the animal. Still, this is more fossil material than is known for any other pterosaur, and it includes both male and female specimens of various age groups and, possibly, species.

Skull and beak
Unlike earlier pterosaurs such as Rhamphorhynchus and Pterodactylus, Pteranodon had toothless beaks, similar to those of modern birds. Pteranodon beaks were made of solid, bony margins that projected from the base of the jaws. The beaks were long, slender, and ended in thin, sharp points. The upper jaw was longer than the lower jaw. The upper jaw was curved upward; while this normally has been attributed only to the upward-curving beak, one specimen (UALVP 24238) has a curvature corresponding with the beak widening towards the tip. While the tip of the beak is not known in this specimen, the level of curvature suggests it would have been extremely long.

The most distinctive characteristic of Pteranodon is its cranial crest. These crests consisted of skull bones (frontals) projecting upward and backward from the skull. The size and shape of these crests varied due to a number of factors, including age, sex, and species. Male Pteranodon sternbergi, the older species of the two described to date (and sometimes placed in the distinct genus Geosternbergia), had a more vertical crest with a broad forward projection, while their descendants, Pteranodon longiceps, evolved a narrower, more backward-projecting crest. Females of both species were smaller and bore small, rounded crests. The crests were probably mainly display structures, though they may have had other functions as well.

Skeleton
Other distinguishing characteristics that set Pteranodon apart from other pterosaurs include narrow neural spines on the vertebrae, plate-like bony ligaments strengthening the vertebrae above the hip, and a relatively short tail in which the last few vertebrae are fused into a long rod. The entire length of the tail was about 3.5% as long as the wingspan, or up to 25 centimetres (9.8 in) in the largest males.

Dinosaurs Live highlight 18: Ankylosaurus

Ankylosaurus (meaning "fused lizard") is a genus of ankylosaurid dinosaur, containing one species, A. magniventris. Fossils of Ankylosaurus are found in geologic formations dating to the very end of the Cretaceous Period (about 66.5–65.5 Ma ago) in western North America.

Although a complete skeleton has not been discovered and several other dinosaurs are represented by more extensive fossil material, Ankylosaurus is often considered the archetypal armored dinosaur. Other ankylosaurids shared its well-known features—the heavily-armored body and massive bony tail club— but Ankylosaurus was the largest known member of the family.

Paleobiology
In comparison with modern land animals the adult Ankylosaurus was very large. Some scientists have estimated a length of 9 meters (30 ft). Another reconstruction suggests a significantly smaller size, at 6.25 m (20.5 ft) long, up to 1.5 m (5 ft) wide and about 1.7 m (5.5 ft) high at the hip. Ankylosaurus may have weighed over 6,000 kilograms (13,000 lb), making it one of the heaviest armored dinosaurs yet discovered. The body shape was low-slung and quite wide. It was quadrupedal, with the hind limbs longer than the forelimbs. Although its feet are still unknown, comparisons with other ankylosaurids suggest Ankylosaurus probably had five toes on each foot. The skull was low and triangular in shape, wider than it was long. The largest known skull measures 64.5 centimeters (25 in) long and 74.5 cm (29 in) wide. Like other ankylosaurs, Ankylosaurus was herbivorous, with small, leaf-shaped teeth suitable for cropping vegetation. These teeth were smaller, relative to the body size, than in any other ankylosaurid species. Ankylosaurus did not share the grinding tooth batteries of the contemporaneous ceratopsid and hadrosaurid dinosaurs, indicating that very little chewing occurred. Bones in the skull and other parts of the body were fused, increasing their strength.

Armour
The most obvious feature of Ankylosaurus is its armour, consisting of massive knobs and plates of bone, known as osteoderms or scutes, embedded in the skin. Osteoderms are also found in the skin of crocodiles, armadillos and some lizards. The bone was probably overlain by a tough, horny layer of keratin. These osteoderms ranged greatly in size, from wide, flat plates to small, round nodules. The plates were aligned in regular horizontal rows down the animal's neck, back, and hips, with the many smaller nodules protecting the areas between the large plates. Smaller plates may have been arranged on the limbs and tail. Compared to the slightly more ancient ankylosaurid Euoplocephalus, the plates of Ankylosaurus were smooth in texture, without the high keels found on the armor of the contemporaneous nodosaurid Edmontonia. A row of flat, triangular spikes may have protruded laterally along each side of the tail. Tough, rounded scales protected the top of the skull, while four large pyramidal horns projected outwards from its rear corners.

Tail Club
The famous tail club of Ankylosaurus was also composed of several large osteoderms, which were fused to the last few tail vertebrae. It was heavy and supported by the last seven tail vertebrae, which interlocked to form a stiff rod at the base of the club. Thick tendons have been preserved, which attached to these vertebrae. These tendons were partially ossified (or bony) and were not very elastic, allowing great force to be transmitted to the end of the tail when it was swung. It seems to have been an active defensive weapon, capable of producing enough of a devastating impact to break the bones of an assailant. A 2009 study showed that large tail knobs could generate sufficient force to break bone during impacts, but average and small knobs could not, and that tail swinging behavior is feasible in ankylosaurids, but it remains unknown whether the tail was used for interspecific defense, intraspecific combat, or both.

Dinosaurs Live highlight 17: Velociraptor

Velociraptor (meaning 'swift seizer') is a genus of dromaeosaurid theropod dinosaur that existed approximately 75 to 71 million years ago during the later part of the Cretaceous Period. Two species are currently recognized, although others have been assigned in the past. The type species is V. mongoliensis; fossils of this species have been discovered in Mongolia. A second species, V. osmolskae, was named in 2008 for skull material from Inner Mongolia, China.

Smaller than other dromaeosaurids like Deinonychus and Achillobator, Velociraptor nevertheless shared many of the same anatomical features. It was a bipedal, feathered carnivore with a long, stiffened tail and an enlarged sickle-shaped claw on each hindfoot, which is thought to have been used to kill its prey. Velociraptor can be distinguished from other dromaeosaurids by its long and low skull, with an upturned snout.

Velociraptor (commonly shortened to 'raptor') is one of the dinosaur genera most familiar to the general public due to its prominent role in the Jurassic Park motion picture series. In the films it was shown with anatomical inaccuracies, including being much larger than it was in reality and without feathers. It is also well known to paleontologists, with over a dozen described fossil skeletons—the most of any dromaeosaurid. One particularly famous specimen preserves a Velociraptor locked in combat with a Protoceratops.

Description
Velociraptor was a mid-sized dromaeosaurid, with adults measuring up to 2.07 m (6.8 ft) long, 0.5 m (1.6 ft) high at the hip, and weighing up to 15 kg (33 lb). The skull, which grew up to 25 cm (9.8 in) long, was uniquely up-curved, concave on the upper surface and convex on the lower. The jaws were lined with 26–28 widely spaced teeth on each side, each more strongly serrated on the back edge than the front—possibly an adaptation that improved its ability to catch and hold fast-moving prey.

Velociraptor, like other dromaeosaurids, had a large manus ('hand') with three strongly curved claws, which were similar in construction and flexibility to the wing bones of modern birds. The second digit was the longest of the three digits present, while the first was shortest. The structure of the carpal (wrist) bones prevented pronation of the wrist and forced the 'hands' to be held with the palmar surface facing inwards (medially), not downwards. The first digit of the foot, as in other theropods, was a small dewclaw. However, whereas most theropods had feet with three digits contacting the ground, dromaeosaurids like Velociraptor walked on only their third and fourth digits. The second digit, for which Velociraptor is most famous, was highly modified and held retracted off of the ground. It bore a relatively large, sickle-shaped claw, typical of dromaeosaurid and troodontid dinosaurs. This enlarged claw, which could be over 6.5 cm (2.6 in) long around its outer edge, was most likely a predatory device used to tear into prey, possibly delivering a fatal blow.

Long bony projections (prezygapophyses) on the upper surfaces of the vertebrae, as well as ossified tendons underneath, stiffened the tail of Velociraptor. The prezygapophyses began on the tenth tail (caudal) vertebra and extended forward to brace four to ten additional vertebrae, depending on position in the tail. The stiffening forced the entire tail to act as a single rod-like unit, preventing vertical motion between vertebrae. However, at least one specimen preserves a series of intact tail vertebrae curved sideways into an S-shape, suggesting that there was considerably more horizontal flexibility. These adaptations of the tail probably provided balance and stability while turning, especially at high speeds.

In 2007, paleontologists reported the discovery of quill knobs on a well-preserved Velociraptor mongoliensis forearm from Mongolia, confirming the presence of feathers in this species. The fact that the ancestors of Velociraptor were feathered and possibly capable of flight long suggested to paleontologists that Velociraptor bore feathers as well, since even flightless birds today retain most of their feathers. These bumps on bird wing bones show where feathers anchor, and their presence on Velociraptor indicate it too had feathers.

Dinosaurs Live highlight 16: Diplodocus

Diplodocus is a genus of diplodocid sauropod dinosaur whose fossils were first discovered in 1877 by S. W. Williston. The generic name, coined by Othniel Charles Marsh in 1878, is a Neo-Latin term derived from Greek διπλόος (diploos) "double" and δοκός (dokos) "beam", in reference to its double-beamed chevron bones located in the underside of the tail. These bones were initially believed to be unique to Diplodocus; however, they have since then been discovered in other members of the diplodocid family and in non-diplodocid sauropods such as Mamenchisaurus.

It lived in what is now western North America at the end of the Jurassic Period. Diplodocus is one of the more common dinosaur fossils found in the Upper Morrison Formation, a sequence of shallow marine and alluvial sediments deposited about 155 to 148 million years ago, in what is now termed the Kimmeridgian and Tithonian stages (Diplodocus itself ranged from about 154 to 150 million years ago). The Morrison Formation records an environment and time dominated by gigantic sauropod dinosaurs such as Camarasaurus, Barosaurus, Apatosaurus and Brachiosaurus.

Diplodocus is among the most easily identifiable dinosaurs, with its classic dinosaur shape, long neck and tail and four sturdy legs. For many years, it was the longest dinosaur known. Its great size may have been a deterrent to the predators Allosaurus and Ceratosaurus: their remains have been found in the same strata, which suggests they coexisted with Diplodocus.

Description
One of the best-known sauropods, Diplodocus was a very large long-necked quadrupedal animal, with a long, whip-like tail. Its forelimbs were slightly shorter than its hind limbs, resulting in a largely horizontal posture. The long-necked, long-tailed animal with four sturdy legs has been mechanically compared with a suspension bridge. In fact, Diplodocus is the longest dinosaur known from a complete skeleton. The partial remains of D. hallorum have increased the estimated length, though not as much as previously thought; when first described in 1991, discoverer David Gillette calculated it may have been up to 54 m (177 ft) long, making it the longest known dinosaur (excluding those known from exceedingly poor remains, such as Amphicoelias). Some weight estimates ranged as high as 113 tons (125 US short tons). This review was based on recent findings that show that the giant tail vertebrae were actually placed further forward on the tail than Gillette originally calculated. The study shows that the complete Diplodocus skeleton at the Carnegie Museum of Natural History in Pittsburgh, Pennsylvania, on which estimates of Seismosaurus were based, had its 13th tail vertebra come from another dinosaur, throwing size estimates for Seismosaurus off by up to 30%. While dinosaurs such as Supersaurus were probably longer, fossil remains of these animals are only fragmentary. Modern mass estimates for Diplodocus (exclusive of D. hallorum) have tended to be in the 10 to 16 tonne (11–17.6 ton) range: 10 tonnes (11 tons); 11.5 tonnes (12.7 tons); 12.7 tonnes (14 tons); and 16 tonnes (17.6 tons).

The skull of Diplodocus was very small, compared with the size of the animal, which could reach up to 35 m (115 ft), of which over 6 m (20 ft) was neck. Diplodocus had small, 'peg'-like teeth that pointed forward and were only present in the anterior sections of the jaws. Its braincase was small. The neck was composed of at least fifteen vertebrae and is now believed to have been generally held parallel to the ground and unable to have been elevated much past horizontal.

Diplodocus had an extremely long tail, composed of about 80 caudal vertebrae, which is almost double the number some of the earlier sauropods had in their tails (such as Shunosaurus with 43), and far more than contemporaneous macronarians had (such as Camarasaurus with 53). There has been speculation as to whether it may have had a defensive or noisemaking (by cracking it like a coachwhip) function. The tail may have served as a counterbalance for the neck. The middle part of the tail had 'double beams' (oddly shaped bones on the underside, which gave Diplodocus its name). They may have provided support for the vertebrae, or perhaps prevented the blood vessels from being crushed if the animal's heavy tail pressed against the ground. These 'double beams' are also seen in some related dinosaurs.

Dinosaurs Live highlight 15: Dimorphodon

Dimorphodon was a genus of medium-sized pterosaur from the early Jurassic Period. It was named by paleontologist Richard Owen in 1859. Dimorphodon means "two-form tooth", derived from Greek δι/di meaning 'two', μορφη/morphe meaning 'shape' and οδων/odon meaning 'tooth', referring to the fact that it had two distinct types of teeth in its jaws - which is comparatively rare among reptiles.

Description
Dimorphodon had a large, bulky skull approximately 22 centimetre in length, whose weight was reduced by large openings separated from each other by thin bony partitions. Its structure, reminiscent of the supporting arches of a bridge, prompted Richard Owen to declare that, in far as achieving great strength from light-weight materials was concerned, no vertebra was more economically constructed; Owen saw the vertebrate skull as a combination of four vertebrae modified from the ideal type of the vertebra. The front of the upper jaw had four or five fang-like teeth followed by an indeterminate number of smaller teeth; the maxilla of all exemplars is damaged at the back. The lower jaw had five longer teeth and thirty to forty tiny, flattened pointed teeth, shaped like a lancet. Many depictions give it a speculative puffin-like 'beak' because of similarities between the two animals' skulls.

The body structure of Dimorphodon displays many "primitive" characters, such as, according to Owen, a very small brain-pan[citation needed] and proportionally short wings. The first phalanx in its flight finger is only slightly longer than its lower arm. The neck was short but strong and flexible and may have had a membraneous pouch on the underside. The vertebrae had pneumatic foramina, openings through which the air sacks could reach the hollow interior. Dimorphodon had an adult body length of 1 metre (3.3 ft) long, with a 1.45 meter (4.6 ft) wingspan.

The tail of Dimorphodon was long and consisted of thirty vertebrae. The first five or six were short and flexible but the remainder gradually increased in length and were stiffened by elongated vertebral processes. The terminal end of the tail may have borne a Rhamphorhynchus-like tail vane, although no soft tissues have yet been found of Dimorphodon to confirm this speculation.

Dinosaurs Live highlight 14: Tyrannosaurus

Tyrannosaurus (meaning "tyrant lizard", from Greek tyrannos (τυράννος) meaning "tyrant," and sauros (σαύρος) meaning "lizard") is a genus of coelurosaurian theropod dinosaur. The species Tyrannosaurus rex (rex meaning "king" in Latin), commonly abbreviated to T-Rex, is a fixture in popular culture. It lived throughout what is now western North America, with a much wider range than other tyrannosaurids. Fossils are found in a variety of rock formations dating to the Maastrichtian age of the upper Cretaceous Period, 67 to 65.5 million years ago. It was among the last non-avian dinosaurs to exist before the Cretaceous–Paleogene extinction event.

Like other tyrannosaurids, Tyrannosaurus was a bipedal carnivore with a massive skull balanced by a long, heavy tail. Relative to the large and powerful hindlimbs, Tyrannosaurus forelimbs were small, though unusually powerful for their size, and bore two clawed digits. Although other theropods rivaled or exceeded Tyrannosaurus rex in size, it was the largest known tyrannosaurid and one of the largest known land predators, measuring up to 12.8 m (42 ft) in length, up to 4 metres (13 ft) tall at the hips, and up to 6.8 metric tons (7.5 short tons) in weight. By far the largest carnivore in its environment, Tyrannosaurus rex may have been an apex predator, preying upon hadrosaurs and ceratopsians, although some experts have suggested it was primarily a scavenger. The debate over Tyrannosaurus as apex predator or scavenger is among the longest running in paleontology.

More than 30 specimens of Tyrannosaurus rex have been identified, some of which are nearly complete skeletons. Soft tissue and proteins have been reported in at least one of these specimens. The abundance of fossil material has allowed significant research into many aspects of its biology, including life history and biomechanics. The feeding habits, physiology and potential speed of Tyrannosaurus rex are a few subjects of debate. Its taxonomy is also controversial, with some scientists considering Tarbosaurus bataar from Asia to represent a second species of Tyrannosaurus and others maintaining Tarbosaurus as a separate genus. Several other genera of North American tyrannosaurids have also been synonymized with Tyrannosaurus.

Description
Tyrannosaurus rex was one of the largest land carnivores of all time; the largest complete specimen, FMNH PR2081 ("Sue"), measured 12.8 metres (42 ft) long, and was 4.0 metres (13.1 ft) tall at the hips. Mass estimates have varied widely over the years, from more than 7.2 metric tons (7.9 short tons), to less than 4.5 metric tons (5.0 short tons), with most modern estimates ranging between 5.4 and 6.8 metric tons (6.0 and 7.5 short tons). Packard et al. (2009) tested dinosaur mass estimation procedures on elephants and concluded that dinosaur estimations are flawed and produce over-estimations; thus, the weight of Tyrannosaurus could be much less than usually estimated. Other estimations have concluded that the largest known Tyrannosaurus specimens had a weight exceeding 9 tonnes.

Although Tyrannosaurus rex was larger than the well known Jurassic theropod Allosaurus, it was slightly smaller than two other Cretaceous carnivores, Spinosaurus and Giganotosaurus.

The neck of Tyrannosaurus rex formed a natural S-shaped curve like that of other theropods, but was short and muscular to support the massive head. The forelimbs had only two clawed fingers, along with an additional small metacarpal representing the remnant of a third digit. In contrast the hind limbs were among the longest in proportion to body size of any theropod. The tail was heavy and long, sometimes containing over forty vertebrae, in order to balance the massive head and torso. To compensate for the immense bulk of the animal, many bones throughout the skeleton were hollow, reducing its weight without significant loss of strength.

Skulls
The largest known Tyrannosaurus rex skulls measure up to 5 feet (1.5 m) in length. Large fenestrae (openings) in the skull reduced weight and provided areas for muscle attachment, as in all carnivorous theropods. But in other respects Tyrannosaurus's skull was significantly different from those of large non-tyrannosauroid theropods. It was extremely wide at the rear but had a narrow snout, allowing unusually good binocular vision. The skull bones were massive and the nasals and some other bones were fused, preventing movement between them; but many were pneumatized (contained a "honeycomb" of tiny air spaces) which may have made the bones more flexible as well as lighter. These and other skull-strengthening features are part of the tyrannosaurid trend towards an increasingly powerful bite, which easily surpassed that of all non-tyrannosaurids. The tip of the upper jaw was U-shaped (most non-tyrannosauroid carnivores had V-shaped upper jaws), which increased the amount of tissue and bone a tyrannosaur could rip out with one bite, although it also increased the stresses on the front teeth.

Teeth
The teeth of Tyrannosaurus rex displayed marked heterodonty (differences in shape). The premaxillary teeth at the front of the upper jaw were closely packed, D-shaped in cross-section, had reinforcing ridges on the rear surface, were incisiform (their tips were chisel-like blades) and curved backwards. The D-shaped cross-section, reinforcing ridges and backwards curve reduced the risk that the teeth would snap when Tyrannosaurus bit and pulled. The remaining teeth were robust, like "lethal bananas" rather than daggers; more widely spaced and also had reinforcing ridges. Those in the upper jaw were larger than those in all but the rear of the lower jaw. The largest found so far is estimated to have been 30 centimetres (12 in) long including the root when the animal was alive, making it the largest tooth of any carnivorous dinosaur yet found.

Arms
When Tyrannosaurus rex was first discovered, the humerus was the only element of the forelimb known. For the initial mounted skeleton as seen by the public in 1915, Osborn substituted longer, three-fingered forelimbs like those of Allosaurus. However, a year earlier, Lawrence Lambe described the short, two-fingered forelimbs of the closely related Gorgosaurus. This strongly suggested that Tyrannosaurus rex had similar forelimbs, but this hypothesis was not confirmed until the first complete Tyrannosaurus rex forelimbs were identified in 1989, belonging to MOR 555 (the "Wankel rex"). The remains of "Sue" also include complete forelimbs. Tyrannosaurus rex arms are very small relative to overall body size, measuring only 1 metre (3.3 ft) long. However, they are not vestigial but instead show large areas for muscle attachment, indicating considerable strength. This was recognized as early as 1906 by Osborn, who speculated that the forelimbs may have been used to grasp a mate during copulation. It has also been suggested that the forelimbs were used to assist the animal in rising from a prone position.[54] Another possibility is that the forelimbs held struggling prey while it was killed by the tyrannosaur's enormous jaws. This hypothesis may be supported by biomechanical analysis.
Tyrannosaurus rex forelimb bones exhibit extremely thick cortical bone, indicating that they were developed to withstand heavy loads. The biceps brachii muscle of a full-grown Tyrannosaurus rex was capable of lifting 199 kilograms (439 lb) by itself; other muscles such as the brachialis would work along with the biceps to make elbow flexion even more powerful. The M. biceps muscle of T. rex was 3.5 times as powerful as the human equivalent. A Tyrannosaurus rex forearm also had a reduced range of motion, with the shoulder and elbow joints allowing only 40 and 45 degrees of motion, respectively. In contrast, the same two joints in Deinonychus allow up to 88 and 130 degrees of motion, respectively, while a human arm can rotate 360 degrees at the shoulder and move through 165 degrees at the elbow. The heavy build of the arm bones, extreme strength of the muscles, and limited range of motion may indicate a system evolved to hold fast despite the stresses of a struggling prey animal. Carpenter and Smith dismissed notions that the forelimbs were useless or that Tyrannosaurus rex was an obligate scavenger.

Dinosaurs Live highlight 13: Styracosaurus

Styracosaurus (meaning "spiked lizard" from the Ancient Greek styrax/στύραξ "spike at the butt-end of a spear-shaft" and sauros/σαῦρος "lizard") was a genus of herbivorous ceratopsian dinosaur from the Cretaceous Period (Campanian stage), about 76.5 to 75.0 million years ago. It had four to six long horns extending from its neck frill, a smaller horn on each of its cheeks, and a single horn protruding from its nose, which may have been up to 60 centimetres (2 ft) long and 15 centimetres (6 in) wide. The function or functions of the horns and frills have been debated for many years.

Styracosaurus was a relatively large dinosaur, reaching lengths of 5.5 metres (18 ft) and weighing nearly 3 tons. It stood about 1.8 meters (6 ft) tall. Styracosaurus possessed four short legs and a bulky body. Its tail was rather short. The skull had a beak and shearing cheek teeth arranged in continuous dental batteries, suggesting that the animal sliced up plants. Like other ceratopsians, this dinosaur may have been a herd animal, traveling in large groups, as suggested by bonebeds.

Named by Lawrence Lambe in 1913, Styracosaurus is a member of the Centrosaurinae. One species, S. albertensis, is currently assigned to Styracosaurus. Other species assigned to the genus have since been reassigned elsewhere.

Individuals of the Styracosaurus genus were approximately 5.5 metres (18 ft) long as adults and weighed around 2.7 tons. The skull was massive, with a large nostril, a tall straight nose horn, and a parietosquasomal frill (a neck frill) crowned with at least four large spikes. Each of the four longest frill spines was comparable in length to the nose horn, at 50 to 55 centimetres long (19.7 to 21.7 in). The nasal horn is estimated at 57 centimeters long (19.7 in) in the type specimen, but the horn is only partially complete. Based on other nasal horn cores from Styracosaurus and Centrosaurus, this horn may have come to a rounded point at around half of that length.

Aside from the large nasal horn and four long frill spikes, the cranial ornamentation was variable. Some individuals had small hook-like projections and knobs at the posterior margin of the frill, similar to but smaller than those in Centrosaurus. Others had less prominent tabs. Some, like the type individual, had a third pair of long frill spikes. Others had much smaller projections, and small points are found on the side margins of some but not all specimens. Modest pyramid-shaped brow horns were present in subadults, but were replaced by pits in adults.[5] Like most ceratopsids, Styracosaurus had large fenestrae (skull openings) in its frill. The front of the mouth had a toothless beak.

The bulky body of Styracosaurus resembled that of a rhinoceros. It had powerful shoulders which may have been useful in intraspecies combat. Styracosaurus had a relatively short tail. Each toe bore a hooflike ungual which was sheathed in horn.

Various limb positions have been proposed for Styracosaurus and ceratopsids in general, including forelegs which were held underneath the body, or, alternatively, held in a sprawling position. The most recent work has put forward an intermediate crouched position as most likely. Paleontologists Gregory Paul and Per Christiansen of the Zoological Museum of the University of Copenhagen in Denmark proposed that large ceratopsians such as Styracosaurus were able to run faster than an elephant, based on possible ceratopsian trackways which did not exhibit signs of sprawling forelimbs.

Dinosaurs Live highlight 12: Gastornis

Gastornis is an extinct genus of large flightless bird that lived during the late Paleocene and Eocene epochs of the Cenozoic. It was named in 1855, after Gaston Planté, who had discovered the first fossils in Argile Plastique formation deposits at Meudon near Paris (France). At that time, Planté (described as a "studious young man full of zeal") was at the start of his academic career, and his remarkable discovery was soon to be overshadowed by his subsequent achievements in physics.

In the 1870s, the famous American paleontologist Edward Drinker Cope discovered another, more complete set of fossils in North America, and named them Diatryma (from Ancient Greek διάτρημα, diatrema, meaning "canoe").

Description
Gastornis parisiensis measured on average 1.75 metres (5.7 ft) tall, but large individuals grew up to 2 metres (6.6 ft) tall. The Gastornis had a remarkably huge beak with a slightly hooked top, which was taken as evidence suggesting that it was carnivorous. Gastornis had large powerful legs, with large, taloned feet, which also were considered in support of the theory that it was a predator.

The plumage of Gastornis is unknown; it is generally depicted with a hair-like covering as in ratites, but this is conjectural. Some fibrous strands recovered from a Green River Formation deposit at Roan Creek, Colorado were initially believed to represent Gastornis feathers and named Diatryma filifera. Subsequent examination showed that they were actually not feathers at all but plant fibers or similar.

Paleoecology
At its time, the environment in which Gastornis lived had large portions of dense forest and a moist semiarid, subtropical or tropical climate. North America and Europe were still rather close, and especially since Greenland was probably then covered with lush woodland and grassland, only narrow straits of a few 100 km would have blocked entirely landbound dispersal of the Gastornis ancestors. While there were large contiguous areas of land in their North American range after the Western Interior Seaway had receded, their European range was an archipelago due to the Alpide orogeny and the high sea levels of the Paleocene-Eocene Thermal Maximum; geographically (but not geologically), it was perhaps roughly similar to today's Indonesia.

Classically, Gastornis has been depicted as predatory. However, with the size of Gastornis legs, the bird would have had to have been more agile to catch fast-moving prey than the fossils suggest it to have been. Consequently, it has been suspected that Gastornis was an ambush hunter and/or used pack hunting techniques to pursue or ambush prey; if Gastornis was a predator, it would have certainly needed some other means of hunting prey through the dense forest.

Alternatively, they may have been predominantly scavengers, omnivores or even herbivores. Indeed, the large beak of Gastornis would have been as well suited for crushing seeds and tearing off vegetation. But it seems excessively strong for a purely vegetarian diet, except in the rather improbable case that huge hard-shelled seeds and nuts formed the main food item of Gastornis. Regardless of what these birds ate, the beak may simply have been used for social display though its presence in all known fossils argues against a sexual display role. These contradicting hypotheses, equivocally supported by the material evidence, make the dietary paleobiology of Gastornis impossible to pinpoint.

Dinosaurs Live highlight 11: Confuciusornis

Confuciusornis is a genus of primitive crow-sized birds from the Early Cretaceous Yixian and Jiufotang Formations of China, dating from 125 to 120 million years ago. Like modern birds, Confuciusornis had a toothless beak, but close relatives of modern birds such as Hesperornis and Ichthyornis were toothed, indicating that the loss of teeth occurred convergently in Confuciusornis and living birds. It is the oldest known bird to have a beak. It was named after the Chinese moral philosopher Confucius (551–479 BCE). Confuciusornis is one of the most abundant vertebrates found in the Yixian Formation, and several hundred complete, articulated specimens have been found.

Description
Confuciusornis was about the size of a modern pigeon, with a wingspan of up to 0.7 meters (2.3 ft), and its body weight has been estimated to have been as much as 1.5 kilograms, or less than 0.2 kilograms. C. feducciai was about a third longer than average specimens of C. sanctus.

Confuciusornis shows a mix of basal and derived traits. It was more "advanced" or derived than Archaeopteryx in possessing a short tail with a pygostyle (a bone formed from a series of short, fused tail vertebrae) and a bony sternum, but more basal or "primitive" than modern birds in retaining large claws on the forelimbs, having a primitive skull with a closed eye-socket, and a relatively small breastbone. At first the number of basal characteristics was exaggerated: Hou assumed in 1995 that a long tail was present and mistook grooves in the jaw bones for small degenerated teeth.

The skull of Confuciusornis was equipped with a pointed toothless beak. It was relatively heavy-built and immobile, incapable of the kinesis of modern birds that can raise the snout relative to the back of the skull. This immobility was caused by the presence of a triradiate postorbital separating the eye-socket from the lower temporal opening, like with more basal theropods, and the premaxillae of the snout reaching all the way to the frontals, forcing the nasals to the sides of the snout.

Fossils of Confuciusornis show that it had an exceptionally large humerus (upper arm bone). Near its shoulder-end this was equipped with a prominent deltopectoral crest. Characteristically this crista deltopectoralis was with Confuciusornis pierced by an oval hole which may have reduced the bone's weight or enlarged the attachment area of the flight miscles. The furcula or wishbone, like that of Archaeopteryx, was a simple curved bar lacking a pointed process at the back, a hypocleidum. The sternum (breastbone) was relatively broad and had a low keel which was raised at the back end. This bony keel may or may not have anchored a larger, cartilaginous, keel for enlarged pectoral muscles. The scapulae (shoulder blades) were fused to the strut-like coracoid bones and may have formed a solid base for the attachment of wing muscles. The orientation of the shoulder joint was sideways, instead of angled upward as in modern birds; this means that Confuciusornis was unable to lift its wings high above its back. According to a study by Phil Senter in 2006, the joint was even pointed largely downwards meaning that the humerus could not be lifted above the horizontal. This would make Confuciusornis incapable of the upstroke required for flapping flight; the same would have been true for Archaeopteryx.

The wrist of Confuciusornis shows fusion, forming a carpometacarpus. The second and third metacarpals were also partially fused but the first was unfused, however, and also the fingers could freely move relative to each other. The second metacarpal, supporting the flight feathers, was very heavily built; its finger carries a small claw. The claw of the first finger to the contrary was very large and curved, that of the third intermediate in size. The formula of the finger phalanges was 2-3-4-0-0.

The pelvis was connected to a sacrum formed by seven sacral vertebrae. The pubis was strongly pointing backwards. The left and right ischia were not fused. The femur was straight; the tibia only slightly longer. The metatarsals of the foot were relatively short and fused to each other and to the lower ankle bones, forming a tarsometatarsus. A rudimentary fifth metatarsal is present. The first metatarsal was attached to the lower shaft of the second and supported a first toe or hallux, pointing to the back. The formula of the toe phalanges was 2-3-4-5-0. The proportions of the toes suggest that they were used for both walking and perching, while the large claws of the thumb and third finger were probably used for climbing.

Feathers
The wing feathers of Confuciusornis were long and modern in appearance. The primary wing feathers of a 0.5 kilogram individual reached 20.7 centimeters in length. The five longest primary feathers (remiges primarii) were more than 3.5 times the length of the hand and relatively longer than those of any living bird, while the secondary feathers of the lower arm were rather short by comparison. Thus, the wing shape was very unlike that of living birds, being long and narrow. The primary feathers were asymmetrical to varying degree, and especially so in the outermost primaries. It is unclear whether the upper arm carried tertiaries. Covert feathers are preserved covering the upper part of the wing feathers in some specimens, and some specimens have preserved the contour feathers of the body. Unlike some more advanced birds, Confuciusornis lacked an alula, or "bastard wing". In modern birds this is formed by feathers anchored to the first digit of the hand, but this digit appears to have been free of feathers and independent of the body of the wing in Confuciusornis. According to Dieter Stefan Peters to compensate for the lack of an alula, the third finger might have formed a separate winglet below the main wing, functioning like the flap of an aircraft.[17] Despite the relatively advanced and long wing feathers, the forearm bones lacked any indication of quill knobs (papillae ulnares), or bony attachment points for the feather ligaments.

Many specimens preserve a pair of long, narrow tail feathers, which grew longer than the entire length of the rest of the body. Unlike the feathers of most modern birds, these feathers were not differentiated into a central quill and barbs for most of their length. Rather, most of the feather formed a ribbon-like sheet, about six millimetres wide. Only at the last one quarter of the feather, towards the rounded tip, does the feather become differentiated into a central shaft with interlocking barbs. Many individuals of Confuciusornis lacked even these two tail feathers, possibly due to sexual dimorphism. The rest of the tail around the pygostyle was covered in short, non-aerodynamic feather tufts similar to the contour feathers of the body, rather than the familiar feather fan of modern bird tails.