Protoceratops (from Greek proto-/πρωτο- "first", cerat-/κερατ- "horn" and -ops/-ωψ "face", meaning "First Horned Face") is a genus of sheep-sized (1.5 to 2 m long) herbivorous ceratopsian dinosaur, from the Upper Cretaceous Period (Campanian stage) of what is now Mongolia. It was a member of the Protoceratopsidae, a group of early horned dinosaurs. Unlike later ceratopsians, however, it was a much smaller creature that lacked well-developed horns and retained some primitive traits not seen in later genera.
Protoceratops had a large neck frill, which may have served to protect the neck, to anchor jaw muscles, to impress other members of the species, or combinations of these functions. Described by Walter Granger and W.K. Gregory in 1923, Protoceratops was initially believed to be an ancestor of the North American ceratopsians. Researchers currently distinguish two species of Protoceratops (P. andrewsi and P. hellenikorhinus), based in part by their respective sizes.
Description
Protoceratops was a quadrupedal dinosaur that was partially characterized by its distinctive neck frill at the back of its skull. The frill itself contained two large parietal fenestra (holes in the frill), while its cheeks had large jugal bones. The exact size and shape of the neck frill varied by individual; some specimens had short, compact frills, while others had frills nearly half the length of the skull. The frill consists mostly of the parietal bone and partially of the squamosal. Some researchers, including Peter Dodson attribute the different sizes and shapes of these bones to sexual dimorphism, as well as the age of the specimen, at the time of death.
Protoceratops was approximately 1.8 meters (6 ft) in length and 0.6 meters (2 ft) high at the shoulder. A fully grown adult would have weighed less than 400 pounds (180 kg). The large numbers of specimens found in high concentration suggest that Protoceratops lived in herds.
Protoceratops was a relatively small dinosaur with a proportionately large skull. Despite being herbivorous, Protoceratops appears to have had muscular jaws capable of a powerful bite. These jaws were packed with dozens of teeth, well suited for chewing tough vegetation. The skull consisted of a massive frontal beak, and four pairs of fenestrae (skull openings). The foremost hole, the naris, was considerably smaller than the nostrils seen in later genera. Protoceratops had large orbits (the holes for its eyes), which measured around 50 millimeters in diameter. Behind the eye was a slightly smaller fenestra, known as the "infratemporal fenestra."
Daily activity patterns
The large eyes of Protoceratops has been suggested as evidence for a nocturnal lifestyle. However, subsequent comparisons between the scleral rings of Protoceratops and modern birds and reptiles have indicated a more cathemeral lifestyle, being active throughout the day during short intervals. This suggests that the fight between Protoceratops and the primarily nocturnal Velociraptor indicated by the fighting specimens may have occurred at twilight or under low-light conditions.
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